2020+. Academic Lead for Public Engagement in the School of Biological Sciences, University of Edinburgh. 2019+. MSc Bioinformatics Programme Director, University of Edinburgh. 2018+. Reader in Bioinformatics, University of Edinburgh. 2016-2018. Lecturer in Bioinformatics, University of Edinburgh. 2010-2015. Lecturer in Biology, University of St Andrews. 2005-2010. Research Councils UK Academic Fellow in Evolutionary and Functional Genomics, University of St Andrews. 2002-2005. Research Fellow, University of Reading. 2001-2002. Developer, Ensembl project, Wellcome Trust Sanger Institute. 1999-2001. Software Engineer and Software Engineer II, Analog Devices Inc.